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Olivia and the others of Fringe division attempt to determine the origin of Peter Bishop, who purports himself as Walter's son who had died as a young boy in the original time line.
The Fringe division is brought to investigate the violent murders of a woman and her live-in boyfriend in her home. They find that she was earlier married to Dr. Malcolm Truss a former Massive Dynamic biochemical researcher, and believe her murder was an attempt to access his aborted research on cell cloning by one of the new shape shifters, Nadine, previously seen in "Neither Here Nor There". While at Massive Dynamic, it is revealed that Nina Sharp raised Olivia and her sister after Olivia killed her stepfather and ran away from Walter's Cortexiphan trials. Walter resents Nina for both this and for her interference that led to young Peter's death. Fringe is unable to locate Truss before Nadine does, and place a manhunt to track him down. Meanwhile, Nadine has tried to convince Truss that she is a patient in need of his cloning research, and coerces him to take her to his unmarked laboratory to synthesis the cure. It's revealed she has a daughter, Haley.

As Fringe division attempts to work out Truss' location, Peter, from his cell, uses the intercom electronics to try to convince Olivia and the others that he knows about shape shifter technology. Under guard, Peter is able to use one of the memory discs previously recovered by Fringe to identify a tracking signal allowing Fringe to set off to Truss' laboratory. Truss has come to discover Nadine's truth after she temporarily loses control and takes the form of his former wife, but under threat of death, continues to synthesize the cure.

Truss finishes the cure just as the FBI and Fringe division surround the warehouse containing the lab. After being given the cure, Nadine attempts to fight off the agents, but appears to have dived into the water below. On recovering a body from the water, they realize that Nadine had taken the place of a fallen agent and has escaped their grasp. Nadine uses a Hermes 3000 typewriter stored in a locker at a Boston train station to type a report informing them of the success of Truss' cure; moments later, the typewriter, on its own, types out further instructions for her.

Walter, after trying to resolve his issues with Peter, returns to Fringe to inform Peter that he had two chances to save his son and let him down both times, he will not try to help again for fear of doing more harm. Meanwhile, Olivia experiences an unexplained time fluctuation.

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